TYPE: Costa Rica. Guanacaste: Liberia-La Cruz, Parque Nacional Guanacaste, Volcán Cacao (northern end of Cordillera de Guanacaste), pressed from cultivation, 12 November 2011, R. L. Dressler 6291 (holotype, USJ; isotypes, USJ, JBL, CR; digital plate of the plant that served as the holotype; photo of a flower from the plant that served as the holotype; photo of a flower from the plant that served as a paratype; photo of the flower from another plant that served as a paratype).